In a post-apocalyptic world where cities are mounted on wheels and consume each other to survive, a young apprentice named Tom Natsworthy finds himself thrown from his home city of London after an unexpected encounter with the mysterious Hester Shaw. As they navigate the treacherous landscape, they uncover a sinister plot that threatens the future of the entire world. Together, they must confront dangerous enemies and forge unlikely alliances to prevent catastrophe, all while grappling with their own personal struggles and evolving relationship.